
Senior Investigator

Education
| 1979 | BA, Biology - Pomona College, Claremont, CA |
| 1984 | Ph.D., Biochemistry and Biophysics - University of California, San Francisco, CA |
Professional Appointments
| 1980 – 1981 | Graduate Student, Program in Cell and Developmental Biology, Harvard Medical School |
| 1981 - 1984 | Graduate Student, Department of Biochemistry and Biophysics University of California, San Francisco (advisor: Prof. Marc Kirschner) |
| 1984 – 1986 | Postdoctoral Fellow, Department of Biochemistry and Biophysics University of California, San Francisco (advisor: Prof. Marc Kirschner) |
| 1986 – 1989 | Assistant Professor of Physiology, Harvard Medical School |
| 1989 – present | Board of Tutors in the Biochemical Sciences, Harvard University |
| 1989 – 1998 | Associate Professor of Cellular and Molecular Physiology |
| 1998 – present | Professor of Cell Biology |
Committee and Advisory Board
| 1989 – present | MD-PhD Admissions Committee, Harvard Medical School |
| 1999 – present | NIH Study Section and Program Project Review Committees |
| 2001 – present | Milton Memorial Fund, Harvard Medical School |
| 2004 – present | Telethon Foundation, Milan, Italy, Scientific Board |
| 2005 – present | Financial Aid Committee, Harvard Medical School |
| 2005 – present | Research Animal Welfare Committee, Harvard Medical School |
Selected Publications

